My apologies, The H52 line did have the boost feature. I was ready to sleep but ZL sent me the reply, guess I didn't check very carefully lol.
About the voltage part, note that ZL is referring specifically to the old SC5 series, and the LEDs used in SC5Fd/c are XM-L2 easywhite, which is 6V. I think this makes sense as the high CRI variation of H52, H502d/c, used Philips LUXEON T instead of XM-L2 easywhite. Compared with these two, the H53c has 95 more lumens on max, and ~10 CRI increase. Note that H502d/c do not support 14500 (but still feature an operating voltage range of 0.7-4.2V) but the SC52d does.
Based on the operating voltage of all variations of the older SC5, I'm guessing that all of them have the same exact circuit that is capable of driving a 6V LED, while dropping 14500 support, so it came down to choices, people either pick the SC52/H52 line for 14500 support, or SC5 if they are looking for good tint & CRI while pumping out impressive lumens (maximum brightness for SC5Fd/c is 375 lumens, 185 higher than H502d/c) but brightness wise SC5 basically rendered the need of a 14500 moot as SC5 can produce 500+ lumens with a standard Eneloop, while SC52/H52 could only crank out 300 lumens with said battery choice.
Back to what ZL said: They probably kept the same driver for SC5c Mk II for the 475lm boost (evident by the no 14500 support), but like they said, the said driver is too large for the H53. Compared to XM-L2 which has a typical forward voltage of 2.85V @ 700mA, XP-L2 is rated at 2.82V @ 1050mA, so ZL probably redesigned the H53's driver (the new fancy user programmable mode groups, remember?) so it could drive the XP-L2 fully. My educated guess is that the combination of SC5's large drive, difference between LEDs and the addition of the user programmable mode groups made ZL to drop 14500 support for the H53 line, which makes sense.
Off topic: Now that I got some sleep (going to get some more), I see ZL mentioning
SC53 in their reply.